Kouphichnium Nopcsa, 1923

DESCRIPTION: Parallel tracks that have an alternating set of impressions: 1. V-shaped or chevron shaped impressions; 2. Digitate impressions with several digits extending from a small impression subperpendicular to the long axis of the trackway. Heteropodous tracks with two primary forms: 1.) 2 Chevron-like series of 4 oval to round holes; 2.) Bifid V-shaped flabellar to bird foot-like imprints with/without median drag marks.

BEHAVIOR(S): Locomotion, repichnia.

ENVIRONMENTAL SETTINGS: Marine to continental freshwater aquatic.

POSSIBLE TRACEMAKERS: Horseshoe crabs.

GEOLOGIC RANGE: Ordovician–recent for marine horseshoe crabs, Devonian–Jurassic for freshwater horseshoe crabs.

ADDITIONAL REFERENCES: (Hasiotis, 2002)

REMARKS: Some Nanopus reidiae (Haubold et al., 2005) may actually be Kouphichnium.